Nos. IAND2.] .-iA'.^ 70.1/1' OF SCOMBER SCOMBER. 257 



to the general tissues of the region, and then continue forward, 

 sometimes as two wholly separate nerves, sometimes as two nerves 

 partly fused, and sometimes as a single nerve. One of these nerves, 

 where there are two of them, or a branch of the single nerve, 

 where there is but one, runs upward and forward, mesial to the 

 terminal part of the ophthalmicus trigemini, and entering the 

 anterior portion of the nasal bone innervates the first organ of the 

 supraorbital canal. This distribution of this branch shows that it 

 contains lateral fibers, and that it must accordingly be derived 

 from the buccalis facialis. Its origin from the parent nerve, inde- 

 pendently of the branches that supply the first five organs of the 

 infraorbital canal, strongly suggests, as already stated in describ- 

 ing the canal, that we have here to do with a separate section of 

 the main infraorbital sensory line, and that that section is the 

 homologue of the anterior section of the line in Aiiiia, that is, of 

 the anterior cross commissure of the latter fish. 



The remaining, distal and terminal portion of the fused nerves, 

 which now belongs entirely to the superior maxillary nerve, forms 

 an important anastomosis with the terminal branches of the oph- 

 thalmic nerves, and then continues forward toward the end of the 

 snout, some of its branches passing internal to the anterior end 

 of the maxillary bone. 



There are thus in Scomber, as in Aiiiia, no motor branches aris- 

 ing from the maxillaris superior. In CJiiuicem Cole describes sev- 

 eral such branches. In that fish the maxillary nerve of his descrip- 

 tions does not closely follow the buccalis facialis, as the maxillaris 

 superior does in Amia and in Scomber. There is, however, in 

 Chimara, according to Cole (No. 15, p. 644), a fairly large bundle 

 of fibers sent from the trunk of the trigeminus to the buccalis, the 

 bundle separating into two parts, one of which comes into relations 

 with the so-called inner buccal nerve, and the other with the outer 

 buccal nerve. This in itself, and, furthermore, both the course of 

 the larger part of the so-called maxillary nerve, and the presence 

 in it of large motor and pharyngeal components, all seem to indi- 

 cate that this nerve of CJiimccra must contain fibers which, in Amia 

 and Scomber, are found in the inferior maxillary nerve. Certain 

 of the motor fibers must represent the branch that in Amia inner- 

 vates the levator maxillse superioris muscles, a nerve which, in 



