296 ALUS. [Vol. ^yVIII. 



to receive two or three strands "aus dem Facialisaste des Tri- 

 geminus," From the ganglion of the united roots two nerves arise. 

 One runs dorsally inside the cranial cavity, is joined by a small 

 branch from the most posterior bundle of the vagus root, and 

 issues from the cranial cavity by a special foramen in the supra- 

 occipital region of the skull, the foramen lying behind the fora- 

 men that transmits the so-called dorsal trigeminus-branch. The 

 other and larger branch from the ganglion of the accessorius 

 Weberi, issues from the cranial cavity through a foramen corre- 

 sponding to the occipital foramina of Amia and Scomber, turns 

 downward and backward, sends a branch to the hypoglossal 

 region, unites with the first spinal nerve, and then enters and is 

 distributed to the pectoral fin. Until the distribution of the 

 branches of this nerve to the associated muscle segments of the 

 trunk, and the distribution of the first free spinal nerve also, are 

 known, it seems useless to attempt a comparison with Amia and 

 Scomber. The supposition that the two spinal roots of the nerve 

 must be the equivalent of the five roots of Scomber, and that the 

 so-called trigeminal root may be a sympathetic one is however 

 obvious, and has already been referred to by me in an earlier 

 work (No. 8). Cole (No. 16) has suggested that the trigeminal 

 root is probably the anterior, trigeminal root of the ramus lat- 

 eralis accessorius of his own descriptions. My reasons for dis- 

 believing this have already been given, and it is to be further- 

 more noted that Haller himself probably gives this anterior root 

 of the lateralis accessorius in the nerve called by him the dorsal 

 trigeminus branch. 



In Lota Goronowitsch shows (No. 32, Fig. 5) the first two 

 spinal nerves each arising by a dorsal and a ventral root, and the 

 two nerves uniting to form a single trunk which is presumably 

 the postvagal nerve of the fish. A nerve called by him the nervus 

 Weberi is said to arise from the ganglion of the facialis, and is 

 said to contain fibers from all the segmental nerves of the tri- 

 gemino-facial complex. Its distribution, so far as given, seems 

 to indicate that it is the ramus lateralis trigemini, or ramus re- 

 currens facialis, of other descriptions of teleosts, and Cole (No. 

 16, p. 166) identifies it as such. It certainly cannot be the 

 homologue of the accessorius Weberi of Haller's descriptions. 



