Il8 ANDREWS. 



be in any case under such control from one centre when the 

 substance of the whole is in ceaseless flux from one to the 

 other of the whole group and in all parts of the units. 



[107] The control in this case must, it seems to me, be in- 

 trinsic to the substance as stick, and not to the substance as 

 organis7n. I think, in short, that one must reasonably believe 

 that the living substance, as far as it can be traced optically, 

 is not only irritable and contractile substance, but that it is 

 already conscious ojganistn. By conscious I mean that more 

 basic form of it which I have above termed sentience. And I 

 think that the phenomena recited already in this paper will of 

 themselves alone bear out to great extent such a conclusion. 

 Sentient, but subject to such direction from organized centres 

 of this same sentience as may be supplied by the nucleus, and 

 other areas of organized differentiation, as the cytoplasmic 

 granules, — I assert the living substance to be. 



The complacency with which the facts on all sides of my 

 investigations lend themselves to a founding of the Metazoa 

 on an ancestral rhizome of Filosa, is reflected in these further 

 characteristics of the Filosa. The Filosa, as I should widen 

 the use of that term, are by far the most numerous and widely 

 distributed so far as we know, for they would include all marine 

 Heliozoa, and the Radiolaria as well as the Foraminifera, in- 

 cluding Gromia and similar forms. As a group they are 

 marked by a tendency to coalesce ; and to multiply the nuclei 

 without actual separation of nucleated areas. As a group they 

 are marked by a plasticity of organization, and show a tend- 

 ency to areal differentiation of structure and a power of adap- 

 tation to general environmental conditions in a number of 

 ways, notably by formation of free-swimming, flagellate, larval, 

 masses. Their differentiation is of the special character which 

 I find to distinguish animal organization ; that is, it tends to 

 emphasis and intensification of irritable and contractile function 

 by organization of the elements upon an ectosarcal basis. 



As a class the Filosa are marked by habits of substance 

 metamorphosis for purposes of perpetuation, such as formation 

 of free-swimming, flagellate larvae, and also of a minute germ- 

 like posterity which are freely motile and suggest sperm. 



