STRUCTUEE AND DIVISION OF TRICHOMONAS 127 



and Robertson ('11) thus describe it for Trichomonas (Tetratric- 

 chomonas?) gallinarum. Kofoid and Swezy ('15) beUeve it is 

 composed of two parts, one of which is a centrosome and the 

 other the basal granule for the fiagella. In the material that I 

 have studied this structure frequently appears to be double, 

 that is, composed of two approximately equal parts, and the 

 posterior flagellum is attached to the anterior moiety, while the 

 chromatic basal rod is connected with the posterior one. Such 

 a condition was also described bj^ Wenyon ('07). Since the 

 three anterior fiagella take the stain so slightly, it is difficult 

 to determine what their relation is to the blepharoplast com- 

 ponents. 



Because the three anterior fiagella do not stain very deeply, 

 they are difficult to make out. This difficulty is often increased 

 by the presence of spirochaetes of similar caliber and staining 

 power and by the fiagella taking a position in contact with, or 

 under, the body. In the drawings thej^ have been omitted when 

 not plainly seen. In all cases in which they could be clearly 

 discerned, they appeared to be of equal length, wavy, and about 

 one-half the length of the body, although sometimes shorter. 

 Figures 2 to 5 and 7 to 15 show the fiagella in their typical 

 condition. 



Hartmann ('10), Wenyon ('07), and Kuczynski ('14, '18) 

 figure these fiagella just as I have found them, but the figures 

 for this species given by Kofoid and Swezy ('15) have the anterior 

 fiagella as long as or longer than the body of the animal. On 

 account of this and other differences, one may be led to suppose 

 that the latter authors were dealing with a different species. 



The posteriorly directed flagellum running as the chromatic 

 margin of the undulating membrane is very much longer than 

 the others, extending the length of the body, making six to eight 

 undulations in its course and projecting posteriorly as a free 

 flagellum as long as the anterior fiagella. This posterior part 

 is similar to the anterior fiagella in caliber and staining power, 

 but the intracytoplasmic portion appears to be much thicker 

 and takes the stain intensely. There is some variation in stain- 

 ability depending on the fixative employed, as will be noted 

 elsewhere. 



