296 FERNANDUS PAYNE 



with those which I have obtained in Gryllotalpa borealis, where 

 a study of a large amount of material leaves no doubt as to what 

 actually takes place. In studying his figures the question arises 

 whether the L-chromosome is really made up of an accessory 

 and a bivalent. In the division of this chromosome Vo'mov 

 says the accessory passes to one pole and the bivalent to the 

 other. This is not what we would expect if this chromosome 

 is a bivalent plus an accessory. In fact we would expect the 



~% >^ B V. 



W&^ D 



I I I 



Text fig. 4 Gryllotalpa vulgaris from Naples. A, B, C, spermatogonial divi- 

 sions, 15 chromosomes; D, first spermatocyte division, eight chromosomes; E, F, 

 G, an L-shaped chromosome. 



bivalent to divide equally, one-half of it passing with the acces- 

 sory to one pole and the other half to the opposite pole. If 

 Voinov's interpretation be correct, we have here a bivalent (by 

 bivalent I mean a first spermatocyte chromosome formed by the 

 conjugation of two spermatogonial chromosomes) which divides 

 in only one maturation division, an unheard of event, so far as 

 I am aware, except in cases of irregularities. Further, in my 

 observations on the Naples form, I find an L-shaped chromosome 

 (fig. 4, E, F, G). Figure 4, F, shows it divided into two equal 



