606 GEORGE T. HARGITT 



tion mitosis the chromosomes which go to the different poles 

 may not be alike in size; there may be a quantitative difference 

 in the daughter chromosome masses. 



These facts would also clearly establish another point, viz., 

 which of the two mitoses was a reducing and which an equa- 

 tional division. Since the synaptic mates have not been exactly 

 alike in some of the chromosomes it is certain that the mitosis 

 in which the division of chromosomes produces some unlike parts 

 must mark the separation of whole chromosomes united during 

 synapsis. And likewise the mitosis resulting in an exact halving 

 of chromosomes indicates a splitting of chromosomes and there- 

 fore an equational division. From the evidence presented the 

 first division, therefore, is a reduction division and the second 

 is the equational. It might be added that in spite of the appar- 

 ent absence of a stage which looks like the usual synapsis stage, 

 the behavior of the chromatic bodies in the maturation mitoses 

 gives evidence that a conjugation has occurred. 



Another point of some interest may be considered, viz., Are 

 the chromosomes of different generations related? This does 

 not involve a morphological continuity, for at the present time 

 such a thing is admitted not to be possible. But do chromosomes 

 of one generation arise from particular chromosomes of a previous 

 generation? If we compare the chromosomes of oogonial mitoses 

 (figs. 5, 6) with the chromosomes of the two maturation mitoses 

 of the oocytes (figures of plate 3) it will appear that there is not 

 a great similarity in form and size of the chromosomes of the 

 different generations. The figures showing the chromosomes in 

 the equatorial plate stages are most favorable for this compari- 

 son. In the oocyte mitoses there are always four large and four 

 small chromosomes. In the oogonia it is not so certain that 

 there are eight smaller and eight larger chromosomes, though 

 there are some which are large and others which are small. 

 The differences in size in the chromosomes of one spindle or of 

 spindles of different cells do not seem to be constant, for an 

 examination of the various figures which show chromosomes will 

 at once make it clear that the chromosomes are not the same 

 relative sizes in every case. While differences are also evident 

 in the shapes of chromosomes the use of this as a criterion of com- 



