222 S. SAGUCHI 



cuticular substance is perforated by canaliculae, filled with fluid; 

 the striae are therefore nothing but the septa of these canalic- 

 ulae viewed in profile. The ciUa pass through the striae and 

 come into connection with the basal corpuscles underneath the 

 cuticle. The basal corpuscles, spherical or oval in form, are 

 here also arranged in linear series running, in most cases, parallel 

 to the shorter diameter of the cell-border. Since it seems that 

 the larger diameter of most epithelial cells is oriented in a direc- 

 tion, perhaps parallel to the axis of the intestine, it is natural 

 that the rows of basal corpuscles all run in a direction, which 

 is at right angle to the axis of the intestine, a condition which, 

 I think, is of an important significance for the function ol ciliated 

 cells in this situation. Under the layer of basal corpuscles fol- 

 lows a comparatively narrow, transparent zone, through which 

 fine fibers, arising from the basal corpuscles, pass downwards. 



The chondriocontes in the cytoplasm are chiefly arranged in 

 a vertical direction and are largely accumulated beneath the 

 transparent zone (figs. 6, 8). They seem not to be floating in 

 the plasma-sap, but to be suspended on or imbedded in the beams 

 of the plasma-network. If a reagent which causes a shrinking 

 of the cell, such as sublimate-mixture, be employed (fig. 7) 

 there appears a sinuous or spiral fiber, stained deeply by iron- 

 haematoxylin, and splitting upwards into several filaments, 

 which end under the transparent zone. With Meves's fluid, 

 on the contrary, no such feature appears (figs. 6, 8). From this 

 it is highly probable that this fiber, found by various investi- 

 gators and termed 'cone of rootlets,' is produced by the act of 

 shrinking, which must result in the cohesion of beams of the 

 plasma -network with the suspended chondriocontes. Centro- 

 somes and mitosis are often met with in ciliated cells from 

 this location. 



4. The middle-intestine of Helix. The constituent elements 

 of the intestinal epithelium of this species are the same as those of 

 Anodonta. The ciliated cells (figs. 9, 15) bear comparatively short 

 cilia, which pass through the cuticle and come into connection 

 with basal corpuscles; these latter are very small granules, so 

 closely applied to one another that they appear, at a certain 



