The Energy of Segmentation 311 



Substituting in this the numerical values obtained for pressures 

 and volume, we find that there has taken place, as a result of fer- 

 tilization and processes subsequent to this, an increase in the 

 energy of the egg of 



7.32 X 1,013,300 X .00000021 cu. cm. =1.567 ergs; 



that, analogously, after the first segmentation, the energy is 



6.53 X 1,013,300 X .00000021 CU. cm. = 1.399 ergs. 



This means, that, as involved in or as identical with the first seg- 

 mentation, there has been a resultant energy decrease, therefore, 

 of .168 ergs; or that it has taken this amount of energy, about ^ 

 of the total increase resulting from fertilization, etc., to bring about 

 this cleavage. 



As bringing about the second segmentation, we find by sub- 

 stitution: (6.53-6.40) X I 013 300 X .000 000 21 =.028ergsof 

 energy to have been involved. 



CONCLUSION 



This completes the computation based on the measurements 

 taken in Experiment VI. It could, of course, also have been made 

 for some of the other experiments, and, had our purpose been to 

 determine as accurately as possible the numerical value of the 

 energy of segmentation, then a large number of both experiments 

 and computations would have been necessary, in order from these 

 to get a mean result. But it has been not the numerical result 

 but rather the method, that is, the practicability, on an experi- 

 mental basis, of applying the "work integral" and so the other 

 equations of which it is a special case, that has seemed the more 

 important and been deemed worthy of emphasis. Thus would 

 I forestall the point of the possible criticism that the numerical 

 results themselves are meager, and that they have been found for 

 only two segmentations on data obtained in one experiment. For, 

 while these are the facts, nevertheless, on the other hand, Experi- 

 ment VI and its results can be regarded as typical of further pos- 

 sibilities, while on the other the limitation to two segmentations 



