Peterson - p. 4 
about 58° in Gecko). This low angle makes sagittal (anterior-posterior) 
movement of the girdle highly effective in increasing the length of the 
stride and versatility of the limb. The glenoid faces more laterally than 
in most anoles (except chlorocyanus and pulchellus); both the recovery and 
girdle muscles are more highly modified for mobility than for any other 
anole (except perhaps twig species), The movement of the girdle observed 
in films of equestris is greater than that in cybotes (comparable film 
observation) or distichus (anatomical inference). 
2. Twig anole behavior (A. occultus, A. valencienni): The preferred 
gait is a walk or slow walk in which two feet always contact the ground, 
Twig anoles are generally slower than the canopy forms. A. valencienni can 
negotiate much finer diameter perches than garmani juveniles of a comparable 
size. The short and evenly proportioned limbs permit the animal to go 
through a full locomotor cycle with all four feet using the same perch. 
Twig animals frequently walk with the body pressed to the perch (crawl); 
they can, however, shift to a straight legged or erect slightly faster 
walking gait, Perches are changed by a short hop (leaps are apparently only 
feasible where the animal can actually drop) or by slow acrobatic maneuvers 
reminescent of Chamaeleo or Phenacosaurus. As in the latter genus the short 
limbs and thus short reach is partially compensated for by axial movement. 
Range of limb movement is at least comparable to the canopy giants. 
Morphology (A. valencienni - incomplete): The girdle is set at 28-30° 
from the sagittal plane, and available data suggest that girdle mobility is 
probably at least as great as in equestris., Glenohumeral morphology includes 
some adaptation related to the short humeral segment and greater emphasis on 
long axis rotation of the humerus, These features parallel chlorocyanus, 
but modification of the protractor musculature for a wider arc of movement 
is much more prevasive and clear cut than in the latter species. 
Twig anoles appear to converge on the anoline genera, Phenacosaurus and 
Polychrus, in morphology and behavior. 
3. Trunk-crown behavior (A. evermanni, A. chlorocyanus, A. grahami); 
These animals generally move faster than the twig forms, but unlike the 
trunk or trunk ground forms do not appear to have a trotting or very fast 
walking gait. They negotiate small perches with greater difficulty than 
valencienni but more readily than equestris. This is perhaps a size effect, 
however, because they cannot move surely and rapidly on perches half their 
own diameter, 
Morphology (A. chlorocyanus): The girdle is set at 370 to the sagittal 
plane, but some of the girdle muscles are poorly adapted to rotate the 
shoulder girdle, The morphology of the clavicular articulations suggests 
that chlorocyanus represents a different phylogenetic "experiment" in 
acquiring girdle mobility. It is highly likely that some girdle mobility is 
present but quite unclear how much the clavicular articulations will permit. 
