TIMOTHY MOERMOND (Department of Zoology, University of Wisconsin, 
Madison, Wisconsin 53706) has set as his problem the mode of habitat 
exploitation by animals and the ways in which the modes of habitat use are 
influenced by the structure of the habitat, In particular, animals must 
move through the habitat to find food items and the patterns of their 
movements will be subject to constraint by the configuration of the habitat. 
Moermond's method of study is timed description of movements and 
postures, In the result, he has been able to show that the frequency of 
types of movement in anoles is consistent within a given species and is an 
adaptation to the specific vegetational matrix in which they live. The 
vegetational matrix is defined by three properties: the diameters of the 
perches, the lengths of the perches, and the distances between perches, 
These properties act in concert to determine the proportion of movements 
between perches (jumps) and the types of movements on perches (crawls or 
runs) of the anoles. By comparing the relative frequencies of movement 
types, three general types of anoles are recognized -- jumpers, crawlers 
and runners - each of which can be identified by characteristic proportions 
of the limbs (see the following table). 
Moermond emphasizes the existence of a gradient of vegetational 
matrices from open formations with tight matrices (grasses and bushes) to 
closed formations with loose matrices (forest). No anole can be maximally 
efficient in every part of the vegetational gradient. It is suggested that 
discontinuities in the gradient would promote behavioral and morphological 
specialization to segments of the gradient, The patterns of movements, 
limb proportions and foraging strategy of a given specialist are all found 
to be correlated and highly predictable from the structure (tightness and 
looseness) of the vegetational structure. 
The tables and figures which follow describe the correlation of anole 
ecomorphs' foraging zones and characteristic movement types. 
The work described above was done at two localities in western Haiti, 
one montane with 7 Anolis species, one lowland with 3 species. Moermond 
has now extended his operations to the two-anole island of Grenada in the 
southern Lesser Antilles. 
The two Grenadian species, A. aeneus and A, richardi, were studied in 
an Acacia wood-lot at Grande Anse, 
The two species differ in sun-shade preference, aeneus, the smaller of 
the two, preferring more open situations, and richardi, the larger species, 
preferring more shaded spots with the juveniles of the two species showing 
the sharpest segregation, Within each species adult males, adult females 
and juveniles occurred in somewhat separate (although overlapping) micro- 
habitats, 
