Peterson - p. 6 
6. Grass behavior (A. pulchellus, A. olssoni): The grass anoles are 
specialists in fast acrobatic maneuvers on small diameter perches. Their 
gait patterns and limb movements resemble the trunk-ground forms, but they 
may also crawl like the twig anoles. They have a very long arc of limb 
movement, but it is more restricted in the anterodorsal quadrant than that 
of twig anoles. 
Morphology (A. pulchellus): A less restricted version of the locking 
mechanism of cybotes is apparently present in pulchellus, but there is 
generally very marked convergence on the canopy forms. This is indicated by 
the 32° orientation of the girdle on the body. The protracting musculature 
reorients as in the canopy forms but does so by different morphological 
pathways indicating functional convergence. The shoulder joint faces more 
laterally but is less open than in the canopy forms. This permits more 
anterior excursion of the humerus but less dorsoventral excursion of the 
limb in these anterior positions, 
A variety of ecological interpretations for the interspecific patterns 
are available, 
1) Naively one might look for a continuous ground to canopy series 
reflecting more specialization for arboreality. But the anole ecomorphs 
represent quite distinct types rather than a continuous series. A. 
distichus and A, pulchellus would be very poor fits in such an interpretation, 
2) Schoener and Williams have suggested that body size differences 
partition food resources and are an important factor in species diversi- 
fication, Body size also affects the range of perches which can be used 
and the kinds of behaviors which are appropriate to shift from one perch to 
another, The size difference of canopy giants and twig anoles clearly 
suggests their different structural habitat and behavior. But in other 
cases it is difficult to interpret the behavioral and morphological data in 
this manner. Size differentiation appears to induce complex rather than 
simple allometric effects, Behavioral and functional strategies are 
adapted not strictly to size but to the relationship among perch size, body 
size and the distance between perches. 
3) The variation in anole structural habitats as viewed by Rand, 
Williams and lately by Moermond is characterized by perch height, perch 
diameter, density of perches and variation in perch orientation, Perch 
diameter correlates with more of the comparative data than any other single 
factor. For example, it predicts the marked convergence of grass and 
canopy forms in features related to girdle and limb mobility. Small perches 
with varying orientations probably provide the major selective force for 
increased mobility. Variation in speed of movement, which is apparently correlated 
with perch elevation, is not as dramatic as that in perch diameter preference, 
But it separates the canopy animals from the trunk or grass forms. Adaptations 
for stability in the proximal forelimb articulations and for power in leaping 
