Gorman - p. 2 
they comment, St. Vincent is "one of the least suitable Lesser Antillean 
banks for such an event, as it is small and not fringed with many islands.") 
On the basis of the new electrophoretic evidence and re-evaluation of 
older data (behavioral and ecological), a totally revised colonization 
sequence is proposed: continent to Grenada —y protorichardi; Grenada to 
St. Vincent —+ by intra-island radiation trinitatis and griseus; St. 
Vincent to St. Lucia —» luciae (before differentiation of trinitatis and 
griseus); St. Lucia to Blanquilla and Bonaire —> blanquillanus and 
bonairensis; St. Vincent to Martinique —» roquet; Martinique to Barbados 
and Grenada independently —} extremus and aeneus. The new argument depends 
not only on genetic similarity as estimated from electrophoresis, behavior 
and other characters but on the hypothesis of strict time-dependence for 
differentiation (e.g. Martinique roquet has six subspecies, so is older than 
Barbados extremus which has none). 
Soulé and Gorman also have investigated "genetic variability" in the 
whole roquet group compared with certain Puerto Rican species. They stress 
"time-divergence' as better predictive of variability than niche width. 
3. The hybridizing anoles of Trinidad. With Licht, Gorman has studied 
the annual reproductive cycles of A. aeneus and A. trinitatis from San 
Fernando, Trinidad, The two species differ markedly. A. trinitatis is 
essentially non-cyclical, A. aeneus ceases reproductive activity in the 
winter months (December-March). If Gorman is correct in assuming that 
Trinidad trinitatis were introduced from Kingstown, St. Vincent, where there 
is never a severe dry season, and A, aeneus comes from a seasonally dry part 
of the Grenada Bank, then on Trinidad the aeneus is preadapted to the 
pronounced dry season, whereas trinitatis probably wastes gametes during the 
winter months. This is the first evidence of an ecological difference 
between the species and may account, in part, for the apparent success of 
aeneus, 
Gorman and Yang, working on the same two species, document a low level 
of introgression between the two species. Several electrophoretic markers 
were utilized that had not been studied previously. Whereas in their 
earlier study all animals were electrophoretically either "pure" aeneus, or 
trinitatis, or Fj hybrids (heterozygous for all markers), they have now 
found evidence for backcrossing. One individual field-identified as a hybrid 
had several loci that were "pure" aeneus, and another individual field— 
identified as aeneus was heterozygous for several loci that implied trinitatis 
in its ancestry. 
4, Reproductive cycles of Puerto Rican anoles. Gorman and Licht have 
a study on female cycles that will appear shortly in ECOLOGY, and a second 
paper by Licht and Gorman has been written dealing with the males. 
Six species of anoles were studied for about 15 months of regular 
sampling from one locality on St. Thomas, and about a half-dozen localities 
on Puerto Rico. 
