198 LABIATA. 
of Boraginee and Labiate are therefore on grounds of all 
analogy of structure, formed on a binary plan. It only 
remains to explain how the anomalous structure arises. 
A bilocular single carpellum may arise either from extreme 
inflection of the dorsal suture, (to which there is in many 
instances a tendency,) or of extreme inflection of the com- 
bined placentze. But from the complete division of the two 
carpella that exists in the above two orders, as well as per- 
haps from the situation of the ovula, we are authorized in con- 
sidering the anomaly as arising from complete inflection of 
the dorsal suture. Although in  Verbenacez, Ehretiacee, 
Cordiacez, and etc., the quadrilocularity of the ovarium arises 
from an opposite cause. There is likewise another cir-, 
cumstance to be mentioned as pointing out the correctness 
of that which we have above stated, this is the almost 
universal basilarity of the styles. "This shews that the lobed 
state of the carpela is more developed towards the 
geometrical apex, a fact in accordance with the structure 
of all syncarpous fruits, The only objection to this exists 
in Cynoglossum, in which thestyle proceeds from the geo- 
metrical, which, is hence the true apex. 
It was perhaps from the consideration of this genus, that 
Brown in his Prod. describes the ovula as pendulous, hence 
shewing his usual extreme acuteness. I find in addition, that : 
this is proved by taking into consideration the situation of the 
raphe. This in Labiate is next the axis, hence the lobes 
of the ovarium are not inverted, but in all Boraginee ex- 
cept Cynoglossum, it is external with regard to the axis, 
proving that the lobes are inverted. It is highly curious, 
thatin such, with one exception, there is an obvious tendency 
in the raphe to revertto its true site, viz., next the axi 
since the foramen in all these is at the lowest point of 
the cavity, while the radicle of the mature seed points tO 
the uppermost. The seed is hence adhering, and in its 
development describes a considerable portion of a circle? 
