II 





diagnostic generic characters given by Szysz) lowicz are that the 

 5 carpels (when all present) are opposite the sepals in Tricuspi- 

 daria and opposite the petals in Dubouzetia (I.e., p. 448) ; and 

 that the seeds of Tricuspidaria have a wing-like strophiole, 

 those of Dubouzetia a spiral strophiole (I.e., p. 450). Lack of 

 material has prevented the verification of the two additional 

 characters just mentioned. Szy^zy lowicz also used the union or 

 non-union of the sepals to distinguish the two genera. He 

 seems to have overlooked Brongniart and Gris's second paper 

 (Bull. Soc. Bot. France, vol. x.. p. 476), as he states in the generic 

 description of Dubouzetia that the ovary is 5-celled (Engl. Bot. 

 Jahrb., vol. vi., p. 453). 



Schumann in 1890 practically followed Szyszylowicz (Engl, und 

 Prantl, Ptianzenfam., vol. iii., pars vi., p. 6), as did also Dalle 

 Torre and Harms in 1901 (Gen. Siphonog., p. 304) ; the latter 

 attribute only one species to Dubouzetia. 



It may be useful to give the more certain external characters 

 which separate Tricuspidaria and Dubouzetia, although it appears 

 doubtful whether they are of generic value. 



Tricusjjidaria.— Sepals more or less united ; petals trifid ; 

 stamens 15-20. 



Dubouzetia. — Sepals free ; petals entire ; stamens 25-35. 



The following account of the anatomy of Tricuspidaria and 

 Dubouzetia has been furnished by Mr. L. A. Boodle, F.L.S. 

 Owing to pressure of work, only one species of each genus could 

 be examined. 



"A comparison of the anatomical structure of Tricuspidaria 

 lanceolate^ Miq., with that of Dubouzetia campanuhtta. Panch , 

 shows a rather close agreement in the more important characters, 

 as well as in a certain number of minor details. 



M The two species agree in the following features. In the leaf : 

 the hairs are uni-cellular, and of similar form ; large crystals of 

 oxalate of lime are present in enlarged cells of the palisade- 

 tissue ; some of the smaller veins are accompanied by scleren- 

 chyina, and are connected with the upper and lower epidermis by 

 sclerenchymatous or slightly thickened cells; in the midrib 

 there is a somewhat broken ring of fibro-vascular tissue, having 

 practically the same form and structure in the two cases. In the 

 stem : the cork cells are tabular ; sclerotic cells are present in the 

 inner part of the primary cortex ; there is a ring composed of 

 fibres and sclerotic cells in the pericycle ; the phloem includes 

 bast-fibres (few), and vertical rows of cells containing crystals ; 

 the wood-fibres have simple pits ; the medullary rays of the wood 

 are one cell in breadth ; the vessels have simple perforations, and 

 sometimes form radial rows ; the portions of the vessel-wall in 

 contact with medullary ray-tissue bear large simple pits ; the cells 

 of the pith are lignitied and of different sizes, and some of them 

 contain crvstals. 



" In describing the points of difference between the two species, 

 " T." will stand for Tricuspidaria lanceolata, "D." for Du- 

 bouzetia campanulata, Walls of upper epidermis undulated (T.) ; 

 straight (D.). Spongv parenchyma denser, and with less chloro- 



