322 Journal of Comparative Neurology. 



The sixth branchial nerve, numbered from behind, arises 

 from that epibranchial ganglion which the N. branchio- 

 gastricus of the vagus enters. Here, as well as with the 

 branchial nerves lying cephalad, it can be established that 

 their most important part is derived from the principal gan- 

 glion, and that the portions derived trom the epibranchial 

 and sympathic ganglia are the smaller. 



The branchial nerve of the glosso-pharyngeus, the seventh 

 from behind, and that of the facialis behave like the foremost 

 one of the vagus. All these nerves divide within the gill 

 arch into the cutaneous and muscular branches, the first of 

 which fuses with an epidermal prominence, the subsequent 

 ganglion praetrematicum. 



In the region of the four anterior epibranchial ganglia this 

 arrangement is destroyed, owing to the stomodasum, and the 

 interpretation of the peripheral nerves present here is diffi- 

 cult. Three nerves come into consideration, namely: (i) A 

 short nerve springing from the fourth epibranchial ganglion 

 (numbered from before), which proceeds behind the N. man- 

 dibularis to the epidermis and ends in a small ganglion 

 (probably the r. mand. ext., Fiirbringer); (2) the N. man- 

 dibularis, which, as a stout fibrillar cord, leaves the second 

 principal ganglion of the trigeminus and derives the greater 

 part of the fibrillae directly from the branchial root; a more 

 slender cord from the third epibranchial ganglion joins this; 

 (3) the N. maxillaris; this arises from the principal ganglion 

 and the second epibranchial ganglion. I must let it remain 

 undetermined whether the first epibranchial likewise has a 

 share in this nerve. The N. maxillaris divides into branches 

 for muscles in the upper lip, for the side wall of the stomo- 

 da'um and for the skin of the lip. A twig proceeding along 

 the skin ends in a ganglion lying close under the epidermis, 

 which behaves entirely like the praitrematic ganglia of the 

 gill region. 



There are thus always present in the region of the tri- 

 geminus small ganglia of a third series, which I might annex 



