Kappers, Teleostean and Selachian Brain. /Q 



their position immediately under the ventricular wall. This is 

 Van Gehuchten's "branche radiculaire interne." Johnston 

 also gives a figure of these motor facialis fibers in the ganoids, 

 while Edinger gave a sketch of the subventricular bundle in 

 Barbus fluviatilis, without, however, giving it a name. In this 

 position this bundle now turns frontally and forms Van Gehuch- 

 ten's "branche radiculaire ascendente." This course it follows 

 for quite a long distance (about half as long as the sensory facialis 

 lying above and kterally of it) and then bends outward near the 

 downward bend of the sensory root and always below the latter. 

 Accordingly, the course of this geniculate motor facialis root is in 

 large measure similar to that of the mammals and man, but differs 

 in that in the latter cases it originates from a nucleus situated 

 lower and its fibers bend up and over the abducens nucleus which 

 is higher up. 



Another part of the motor facialis fibers do not take the dorsal 

 course, but after partial decussation take a latero-ventral direc- 

 tion through and immediately beneath the tr. descendens N. V. 

 They gather latero-ventrally of this tract (Figs, xcviii, xcix, Plate 

 VII) and after having followed it in a frontal direction for a short 

 distance they leave the brain together with the upper geniculate 

 facialis root. Of these fibers some originate more ventrally than 

 the other facialis fibers in a cell group which, at least in Lophius, 

 is situated at about the same level as that of the abducens (Fig. 

 Ixxvii, Plate V). Their' course may easily be followed and has 

 also been given by Haller and Johnston. Perhaps this is also 

 the reason why Van Gehuchten, who describes only the most 

 dorsal motor root, describes "prolongements protoplasmatiques" 

 between the nucleus of this root and the tr. descendens nervi quinti, 

 which he considers to have the function of short reflex tract 

 between the sensory trigeminus and the motor facialis. Although 

 I must admit the possible existence of such a reflex tract, yet I can 

 state for certain that these "protoplasmatic prolongations" are 

 motor roots of the facialis going through the descending fibers of 

 the trigeminus. 



The sensory facialis root in Gadus goes laterally of and above 

 the wall of the ventricle, and, as has long been known, arises from 

 the most anterior part of the lobi vago-glossopharyngeales. 

 According to Haller, C. J. Herrick and Johnston the root 

 contains only sensory fibers, which Kingsbury contradicts and 



