8o yournal of Comparative Neurology and Psychology. 



considers it to have also a viscero-motor function. I cannot 

 confirm Kingsbury's statement on this question, although it 

 accords with Kohnstamm's discovery that in the higher verte- 

 brates, after resection of the chorda, large (secretory?) ganglion 

 cells degenerate in the taste nucleus. 



The course of the sensory facialis fibers caudad has been 

 described in the highest vertebrates for the pars intermedia 

 (Wrisbergi) of this nerve which in these forms also ends in the 

 lobus glossopharyngei (His, Duval, Edinger). That this root 

 originates from the glossopharyngeus region might be a motive 

 for not considering it a genuine facialis root but a part of the 

 glossopharyngeus which runs out with the facialis. In this 

 connection it is of great importance to determine what connection 

 there is between the terminations of these sensory facialis fibers 

 and the sensory glossopharyngeus fibers, in other words whether 

 their agreement in central origin is borne out by their peripheral 

 ramifications. 



Investigations on their peripheral course have been made first 

 by Cole (Gadus) and by C. J. Herrick (Menidia), who both 

 drew the conclusion that from the ganglion geniculi sensory 

 facialis fibers originate which by reason of their course and mode 

 of termination form the rudiment of the chorda tympani of man. 

 That this peripheral relation does exist is positively proved by this 

 fact together with the further fact that the chorda provides the 

 anterior and the glossopharyngeus the posterior part of the tongue 

 with taste fibers. Jacobson's anastomosis of Gadus, according 

 to Cole, must also be considered as another argument for the 

 close relation between the glossopharyngeus and facialis. 



Nervus ocfavus and nervi laterales. Before describing these I 

 must take up an important question which has been prominent 

 in the literature for many years. The earlier investigators of the 

 selachian brain, viz., Rohon, Sanders and Viault, described in 

 the sharks and rays a considerably elevated part of the gray sub- 

 stance at the most anterior latero-dorsal region of the medulla 

 oblongata, in which originated a large medullated root which was 

 evidently sensory from its mode of origin and termination. This 

 root was considered by them as belonging to the trigeminus and 

 for a long time was described as such. By reason of this concep- 

 tion the excrescence of the octavus field, where the lobus is really 

 situated, was called the lohus trigemini. Mayser, to whom we 



