256 "Journal of Comparative Neurology and Psychology. 



other source can be traced into the vicinity of the lateral line organs 

 in question, the communicating nerve from the r. buccahs to the 

 palatine must be the only path by which lateralis fibers can reach 

 these organs. This is strong indirect evidence to show that the 

 communicating nerve carries lateralis fibers, but it does not prove 

 that it is exclusively lateraHs in composition. The possibihty 

 remains that it is composed partly of general cutaneous fibers. 

 However, it would not seem reasonable for general cutaneous 

 fibers to travel this round-about road to a region which is already, 

 by most direct route, profusely supplied with general cutaneous 

 innervationby the mesial terminal ramus of the ophthalmicus pro- 

 fundus. Both indirect and direct evidence, therefore, is very strong 

 to prove that the communicating nerve between the r. buccalis 

 and the palatine nerve belongs to the acustico-laterahs system. 



J. The Communis Component of the Facial. — Beyond the 

 descriptions of the terminal branches of the r. palatinus, already 

 given, there is nothing particularly noteworthy regarding this 

 nerve in Triton as compared with Amblystoma. It arises from 

 the cephalic portion of the geniculate ganglion, from which the 

 fibers of the r. alveolaris pass directly laterad with the hyoman- 

 dibular trunk. The nerve called palatinus caudalis in Amblys- 

 toma occurs also in Triton. It varies considerably in its point of 

 origin, sometimes coming out of the ganglion almost at the base 

 of the r. palatinus, sometimes about half way through the facial 

 canal leaving the hyomandibular trunk through a special foramen. 

 In one case it seemed to be represented by fibers which leave the 

 communis component of the hyomandibular just outside of the 

 facial canal. My preparations show conclusively that this is a 

 communis nerve from the geniculate ganglion, and that it anasto- 

 moses with a nerve of about the same dimensions from the glosso- 

 pharyngeus. The fibers of the two nerves pass cephalad together 

 from the point of anastomosis. They can be traced cephalad in 

 the roof of the mouth as far as the central level of the eye. This 

 is the true Jacobson's anastomosis, and should not be confused 

 with the other connective between the vagus and facial. 



There is one peculiarity of the r. alveolaris: It receives no fibers 

 from the glossopharyngeus, such as were described in Amblys- 

 toma. There is a large branch of constant occurrence which 

 arises near the angle of the jaw and turns mesad to the epithelium 

 between the hyoid and mandibular arch. 



