CocuHiLL, Fifth Nerve in Amphibia. 49 
nerves arise wholly separate from any branch of the Gasserian 
ganglion, and are, therefore, in their proximal portions purely 
lateral line nerves. In some larval heads I have examined, 
these nerves are more obscure in their relations than I find them 
in the adult. 
One of these lateral line nerves is the superficial ophthalmic 
Vil (Fig. 4, os). It passes laterad out of the cranium in com- 
pany with other branches from the Gasserian and lateral line 
ganglion, between the muscles masseter and temporalis, then 
meso-cephalad subcutaneously mesal of the eye. It innervates 
the supra-orbital lateral line organs. It is intimately associated 
with a twig from the opthalmicus profundus V over the nasal 
septum, but inno case have I found a true anastomosis between 
these nerves. The ophthalmicus superficialis VII is, therefore, 
a purely lateral line nerve throughout. 
The other nerve issuing from the lateral line ganglion in 
close relation with the superficial ophthalmic is the ramus buc- 
calis VII. It at first follows a course ventral of the superficial 
ophthalmic VII, and as it passes out of the cranium fuses with 
a general cutaneous nerve from the Gasserian ganglion (Fig. 4, 
6). The resulting nerve will be discussed blow. . 
The dorso-lateral portion of the Gasserian ganglion gives 
rise to the ramus mandibularis V, which, farther than its origin 
from the ganglion and its relation to the nerve which joins the 
buccal VII, has no bearing on the present discussion. 
From the ventral portion of the Gasserian ganglion the 
ophthalmicus profundus V passes directly cephalad, and, after 
giving off numerous branches of no special significance here, 
divides in the region of the optic nerve into three terminal 
branches. These nerves I designate, for purposes of this dis- 
cussion, as the lateral, mesal and ventral terminal branches of 
the opthalmicus profundus V (Fig. 1, o/, om, ov). 
The mesal terminal branch of the ophthalmicus profundus 
(Fig. 1, om) inclines mesad to a position dorsal of the muscle 
obliquus superior and mesal of the olfactory epithelium, near 
the dorsal wall of the olfactory capsule. Within the capsule 
it gives off several twigs which penetrate the roof of the cap- 
