CocHitL, f2fth Nerve in Amphibia. 55 
being the case, we have in Amphibia an infra-orbital nerve of 
acustico-lateral and general cutaneous composition. 
The relations of this infra-orbital nerve in the adult Rana 
seem to be but partially known. The acustico-lateral component 
degenerates with the disappearance of the lateral line organs. 
The general cutaneous component probably persists as the rami 
zygomatico-temporales and palpebrales inferioris as described 
by Gaupp. ‘The relation which the nerve holds to the maxillo- 
mandibularis in older tadpoles suggests the possibility of a 
more extensive fusion of the nerve with that trunk so that the 
at one time independent twig might later become a ramus of 
the maxillaris. It is quite certain, at any rate, that the region 
innervated in the adult by the rami mentioned is not touched 
by the maxillaris of the larva but is innervated by the ‘‘acces- 
sory ramus.” 
Although the relations existing in other Amphibia between 
the ophthalmicus profundus, the maxillaris, and the palatinus 
VII have received but incidental notice by anatomists generally, 
a brief review of the subject may be helpful. 
In Siredon pisciformis (larva of Amblystoma) FiscHEr de- 
scribes the ventral terminal branch of the ophthalmicus pro- 
fundus (his nasalis) as issuing from the lateral terminal branch 
of that nerve. He finds the anastomosis of the ventral branch 
with the palatinus VII, and describes the nerve resulting from 
that anastomosis as passing cephalad over the vomer essentially 
as I find the mesal nerve derived from the double anastomosis. 
The anastomosing nerves which pass laterad immediately cau- 
dad of the internal nares escaped his notice. The irfra-orbital 
nerve, which I have described as composed of acustico-lateral 
and general cutaneous components, Fischer calls the maxillaris 
superior of the trigeminus. The peripheral distribution he as- 
signs to this nerve fulfils all the conditions of both components. 
In Amblystoma punctatum, Herrick describes the ophthal- 
micus profundus as forming into three terminal branches, and 
mentions the anastomosis with the palatinus VII as a ‘‘broad 
commissure.” His figures suggest that he saw the two nerves, 
the mesal and lateral, derived from this anastomosis, but he has 
