Herrick, Werves of Siluroid Fishes. 193 
ous branches for the adjacent parts of the wide palate and its 
taste buds, some branches running out intv the mucous lining 
of the suspensorium (hyomandibulare, cartilaginous symplectic 
and quadrate) and of the proximal part of the hyoid arch. 
This whole region is freely supplied with taste buds which are 
innervated from these nerves. The middle and ventral parts of 
the hyoid arch are also supplied with taste buds, but they are 
innervated from a different source ; viz. by a twig of the lingual 
branch of the post-trematic branch of the glossopharyngeus, 
which turns caudad from the point of union of the hyoid arch 
with the copula and follows the dorsal surface of the ceratohyal 
back as far as the region supplied by the nerve now under con- 
sideration. The lining of the mandible is freely supplied with 
taste buds, especially toward the distal end of the ramus. But 
none of these are supplied from the facialis, but from a branch 
of the r. mandibularis V which early becomes detached from 
that nerve and finally terminates in the lower breathing valve. 
The posterior palatine branch of the facialis in Ameiurus 
as compared with the corresponding nerve of other fishes pre- 
sents two features which are apparently abnormal; viz. (1) its 
origin is far removed from that of the r. palatinus, and (2) it 
runs down caudad of the pseudobranch, instead of cephalad of 
that organ. Its position is, however, really perfectly typical, 
the other two organs having been displaced cephalad. The r. 
palatinus arises from the infra-orbital trunk much farther ceph- 
alad than in teleosts in general, and this can only be regarded 
as a secondary modification. The pseudobranch (first described 
by McKeEwnzig, ’84, p. 426, whose description I confirm) is in a 
very rudimentary condition, a mere rete mirabile on the inter- 
nal carotid artery (Fig. 3, psdv.). That it has suffered very 
great secondary displacement cephalad is obvious, its anterior 
extremity actually lying under the foramen by which the optic 
nerve leaves the cranium. The r. palatinus runs along its outer 
face and if it receives any cerebro-spinal nerve supply (which I 
consider improbable), it would doubtless be from this latter 
nerve. 
The abnormal position of the nerve which I here term the 
