208 JouRNAL OF COMPARATIVE NEUROLOGY. 
_gans in the siluroid fishes described by Porrarp (’92). In 
these cases also there are cutaneous branches not destined for 
canal organs; but he does not indicate their composition. Of 
course, in enumerating this branch with the glossopharyn- 
geus we must not lose sight of the fact that this is for conve- 
nience of topographical description merely. It has no real 
morphological connection with that nerve, being a branch of 
the r. lateralis vagi. 
The conditions above described were found also in a sec- 
ond specimen and may be regarded as typical for this species, 
though on the opposite side of the specimen plotted this later- 
alis twig separates from the IX root intra-cranially and passes 
out through the vagus foramen, the other relations being as 
above described. 
The remainder of the glossopharyngeus runs forward out- 
side the cranium to its ganglion in the usual way. The details 
of the peripheral relations have not been fully worked out. 
The post-trematic ramus is very large. I have not succeeded 
in demonstrating a pre-trematic ramus, nor does WRIGHT men- 
tion it in his account of A. catus, and both Stannius (’49, p. 76) 
and JuGE (’99, p. 103) deny its presence in Silurus. The pala- 
tine ramus is distinct and can be followed as far forward on the 
roof of the mouth as the level of the chief foramen of the V+ 
VII nerve complex, where it supplies large taste buds of the 
palate. No JAcogpson’s anastomosis was discovered; compare 
the discussion of the r. palatinus posterior, above. 
III. THE VAGUS NERVE. 
The vagus roots, motor, communis, lateralis and general 
cutaneous, are arranged in the typical way, the motor root 
arising in several distinct strands ventrally of the spinal V tract 
and the communis root passing out dorsally of this tract. The 
general cutaneous root is quite large and joins the communis 
root as the latter passes over the spinal V tract. The general 
cutaneous fibers derived from the vagus follow the spinal V 
tract for only a very short distance, soon turning inward to 
terminate in the substantia gelatinosa mesially of the tract. 
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