GENETIC STUDIES ON FLOUR-MOTH 421 



divided while the distal part is normal. Failure of the maxillae 

 to unite is apparently due to malformation. 



Figure g shows a condition in which the palpi are very small, 

 the tongue is cleft to the base and a short distance from the base 

 the elements diverge laterally. In figure h the palpi are normal; 

 the maxillae are separated distally. Occasionally a moth fails 

 to shed the pupal covering of the head. It may otherwise be 

 quite normal. Figure i illustrates this, showing a ventral view 

 in which also the maxillae are separated and coiled up at the 

 sides. 



Other variations are swellings on the tongue or antennae, 

 straight tongue and scaleless areas on wings. 



2. Analysis of original stocks. Summaries of the earlier results 

 will be given in order to show the origin of the variations studied. 



a. Bussey stock. Defective mouth parts. Nine non-virgin 

 type females from the Bussey stock, when isolated, produced 

 841 type, 448 males and 393 females; 6 with defective palpi, 2 

 males and 4 females; 5 with deformed tongue, 2 males and 3 

 females; and one male with defective genitalia. Defects of palpi, 

 tongue, and male genitalia, therefore, occur in the Bussey stock. 



Twenty virgin females from the Bussey stock were paired with 

 males, either from the Bussey, the Lowell, or the Washington 

 stock. They produced 473 type, 255 males and 218 females; 

 3 with defective palpi, 1 male and 2 females, and 1 female with 

 cleft tongue. Each of these abnormalities occurred among the 

 offspring of a different mating, so that no significant Mendelian 

 ratio appeared. 



b. Lowell stock. Defective mouth parts and sooty base. Ten 

 non-virgin type females from the Lowell stock, w^hen isolated, 

 produced 917 type, 452 males and 465 females; 24 with defective 

 palpi, 13 males and 11 females; 7 with deformed tongue, 4 males 

 and 3 females. Defects in palpi and tongue therefore occur in 

 the Lowell stock as in the Bussey stock. 



A pair of type produced 55 type, 25 males and 30 females; 

 and 19 with cleft tongue, 9 males and 10 females. Another 

 similar mating produced 27 type, 17 males and 10 females, and 

 8 with cleft tongue, 4 males and 4 females. This amounts to 



