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WILLIAM A. KEPNER AND ARNOLD RICH 



as indicated at (F). Immediately the oral end opened and 

 closed in an indifferent manner, so that food, which lay near the 

 mouth, was not ingested. No peristaltic waves arose in the 

 aboral part of the organ. Finally, in a third specimen (fig. 3), 

 we had a case in which the freed proboscis was quite active 

 when complete (G) ; but there were evident no longer any of 

 the movements which play a role in the ingestion of food when 

 the oral fourth of the proboscis was removed (H). A marked 

 variation from the reaction of the above incomplete proboscis is 



C 



Fig. 2 A, entire animal; B, dorsal ganglia and anterior portion of body re- 

 moved; C, posterior portion of body cut away; D, proboscis severed through base, 

 sphincter removed, proboscis then swam from sheath and remained inactive ;£", 

 free proboscis; F, proboscis severed transversely, oral end opened and closed in 

 an indifferent manner. 



that of the specimen shown or represented in our figure 4. 

 Here the sphincter (D) was cut away, and though the movements 

 of the remainder of the proboscis were not normal (more spas- 

 modic) , still the proboscis mouth explored for food and ate. The 

 larger part of this proboscis swam about, and in coming by its 

 own detached sphincter accepted the latter by greedily ingesting 

 it.. Unless, therefore, the proboscis be complete enough to per- 

 mit of all of the three coordinated movements being effected, 

 no two and usually no one of this set of movements will be well 

 carried out. 



