562 SHINICHI MATSUMOTO 



Naturalh^ the question arises, whether the epithelial movement 

 on the tissue, which is so frequently to be seen in the fluid me- 

 dium, is chemotactic or thigmotactic in nature, It might even 

 be that the type of movement is due both to mechanical and to 

 chemical influences acting simultaneously. However, the epi- 

 thelium is able to move with practically the same velocity both 

 on a substratum where the covering epithelium has been simply 



Fig. 9. Expermient 239. Corneal piece cultivated in serum with pith; three 

 days after explanation. Note the epithelium clinging to the piece of pith (p) 

 moving out of the cut end (/) of cornea (c). X 98. 



scraped off and on one where the underlying tissue has been 

 killed by heating. The movement in the latter cases cannot, 

 therefore, be dependent upon chemotactic influences from the liv- 

 ing tissue. The same is true of the movement taking place on 

 the surface of the cartilaginous plate of the sclerotic coat, pre- 

 viously killed by boiling. 



Furthermore, the epithelium is able to move on the surface 

 of glass, on a celloidin film, and also on such fiber-like supports 



