540 P. W. WHITING 
University of Pennsylvania for the expense of the experiments. 
I also wish to thank the Zoological Society of Philadelphia for 
the opportunity of crossing my cats with the Caffre cat. 
Before presenting the data and discussing the inheritance of 
the various characters in detail it may be well to name and to 
define briefly the factorial differences involved. 
The ticking or agouti series contains at least two and probably 
more different factors. Black, non-agouti, or uniformity, a, is a 
recessive allelomorph to agouti, A. An extreme amount of 
ticking is frequently observed, and this represents, in all proba- 
bility, a third allelomorph in the series. Results are not con- 
clusive, but are consistent with such a hypothesis. This ex- 
treme ticking may tentatively be called A’. 
The banding or tabby series contains at least two and probably 
three different allelomorphs. They may be called lined, B’, 
striped, B, and blotched, 6. The allelomorphism of these is 
again merely tentatively assumed, but I believe there is good 
reason to consider them as a triple allelomorphic series. The 
agouti and the banding factors will be discussed fully later in 
this paper. It may merely be stated here that all cats are 
either lined, striped, or blotched and that there are no in- 
tergrades. The character of the banding is, however, easily 
recognized only in the presence of ticking. In the black or 
non-agouti series banding exists as ‘ghost patterns’ recognizable 
usually but not always in the fur of kittens, and only occasionally 
distinguishable in full-grown cats. 
Intense pigmentation, M, is a simple dominant over dilute 
or maltese, mm. 
White, W, is a simple dominant over color, w. 
Yellow, Y, is allelomorphie with black, y. This locus is sex- 
linked and shows the male to be digametic. The heterozygous 
female is usually yellow-spotted or tortoiseshell, but may range 
in color to solid black or solid yellow. 
White-spotting is very irregular and probably depends upon 
many factors. It will not be considered in detail in the present 
paper. It appears to segregate independently of the factors above 
mentioned. 
