No. 3.] ALLOLOBOPHORA FOETIDA. 499 
becoming individualized and the mass forming an approximately 
homogeneous fluid, whereas the spermato- 
phores do not disintegrate in water, even 
after twenty-four hours’ immersion. This 
suggests that the spermatozoa, after leav- 
ing the spermathecae, come in contact 
with some adhesive substance which welds 
them into masses and confines them within 
the area which is occupied later by the 
cocoon. Removing the slime-tube from 
worms with partly formed cocoons, I found on the surface of 
segments 9g to II, inclusive, tiny opaquely white specks (some- 
times as many as nine) apparently issuing from integumental 
orifices. These, like the spermatophores, did not disintegrate 
in water, and when stained and mounted they showed a very 
definite granular structure (text Fig. 3). 
The glands which secrete this substance can be seen on 
dissecting the worm from the ventral surface and removing 
everything but the spermathecae, leaving the inner surface of 
the integument exposed. On segments 9 to 11, inclusive, are 
tiny opaquely white swellings. If these are pricked with a 
needle, an opaquely white substance can be pressed from them, 
which, when stained and mounted, entirely resembles that 
described above as found on the exterior of the same segments. 
In some worms these tiny swellings are quite numerous, one 
being close to the stem of each spermatheca and at least four 
in each of the three segments, these being distributed in the 
center of the segment as well as close to the dissepiments. 
If the center, more dense part, of a spermatophore is crushed 
on a slide, stained and mounted, we find the same granular 
secretion as in the above-mentioned integumental glands of seg- 
ments 9 to 11, and on their exterior surface. Thus it appears 
to be demonstrated that the spermatophores of this worm 
are formed by the spermatozoa aggregating around the gran- 
ular— probably nutritive — substance secreted by these glands. 
Archoplasm.— The appearance of the archoplasm (Foot (6) ) 
in the living egg indicates that it is at least semi-fluid, and this 
interpretation is in accord with the history of the archoplasm, 
