502 FOOT. [Vou. XIV. 
of the second spindle, each of the eleven chromosomes is repre- 
sented by two bent, straight, or contracted rods,! this longi- 
tudinal division resembling that of the twenty-two chromosomes 
of the cleavage spindle. The phenomena in this egg offer no 
evidence in support of Hiacker’s (g) last theory of reduction ; 
for to meet the demands of this theory there must be some 
evidence that the two parts of each chromosome of the second 
spindle are individually the same two parts of the tetrad that 
remained in the egg after the first division. This cannot be 
asserted, for the reason that between the first and second 
divisions we have a semi-resting stage,” that is, the chromo- 
somes take the first steps towards a typical resting stage, 
forming vesicles extirely similar to those formed after the 
second division and after the first cleavage. There are eleven 
vesicles. This indicates that two parts of each tetrad of the 
first spindle have fused into one vesicle, and the individuality 
of each is thus lost. The process of the reorganization of 
these vesicles into the chromosomes of the second spindle 
must occur very rapidly ; for none of my preparations show 
this step in the development. - 
This egg appears to show only two methods of reduction, — 
first, the above-mentioned number reduction, and second, a 
mass reduction. The latter is proved by the omission of 
a typical resting stage and by the amount of chromatin in the 
second spindle as compared with that in the first. In text Fig. 4 
are represented some of the forms shown by the tetrads of the 
first maturation spindle. I am unable at present to give any 
data regarding the formation of these tetrads, as I have not 
1 For figures representing these chromosomes, see Foot (5), Fig. 3; (6), Fig. 10; 
(8), Fig. 4. 
For polar view of the tetrads (5), Fig. 4, and Fig. 5 for polar view of second 
maturation spindle. At the time these last two figures were drawn, I was unable 
to interpret the apparent tetrads often found in the second maturation spindle. 
The following summer, in preparations differently fixed, the rods showed a suc- 
cession of bead-like formations (6 or more), this indicating that the swelling of 
each rod into two or several of these bead-like formations is possibly due to the 
fixative or some other artificial factor. 
21 designate this as a semi-resting stage, for the reason that the chromatin 
lost by the last division is not replaced by growth, this being an essential feature 
of the typical resting stage. : 
