NERVE ENDORGANS IN THE EAR 299 
From the above it appears, then, that the macula neglecta 
Retzii, which comes to lie in the floor of the utriculus, arises from 
the neuroepithelium of the pars inferior, as was first established by 
Fleissig in the case of Gecko; but, as stated above, I am unable 
to demonstrate in my material any progressively developing 
endorgan which could represent the macula neglecta Sarasini. 
Alexander has suggested that in the embryo of Echidna the 
tall epithelium at the mouth of the ductus endolymphaticus may 
represent the vestige of the Amphibian macula neglecta Sarasini. 
This tall epithelium, which is continuous with the neuroepithelium 
of the medial utricular wall, Fleissig has also observed in the em- 
bryo of Gecko, but his interpretation is a totally different one, 
for he does not consider it remarkable that the mouth of the 
ductus endolymphaticus, which is still in active growth, should 
possess tall epithelium where it passes suddenly into the neuro- 
epithelial anlage of the medial utricular wall. 
In conclusion I desire to record the observation that the three 
semicircular canals of Trigonocephalus japonicus do not develop 
synchronously, the medial and posterior canals anticipating the 
lateral canal in their development. 
SUMMARY 
1. The macula neglecta arises directly from the macula partis 
inferioris. 
2. The occurrence of two maculae neglectae is not to be 
observed in my material: while the macula neglecta Retzii is well 
developed, there does not form a persistent macula Sarasini nor 
does this endorgan even develop temporarily as in Gecko (Fleis- 
sig). 
3. The anterior and the posterior semicircular canals are 
separated off much earlier than the lateral canal. 
Kyoto, Sept. 15, 1914. 
