396 SYDNEY E. JOHNSON 
are more in evidence. In the posterior ganglia (9 and 10) no 
spirals could be found and only a few broken sections of pregan- 
glionie fibers. The spinal cord of this specimen was found to 
have been destroyed as far forward as the third vertebra. 
This experiment appeared to eliminate the possibility of incom- 
plete degeneration, but it also left the origin of a large number of 
spiral fibers to be accounted for. Two possibilities presented 
themselves: 1) cerebrospinal fibers could still reach the sympa- 
Fig. 10 Small section of 5th or 6th ganglion of a specimen injected with 
methylene blue three months after destruction of the spinal cord. Many spirals 
and pericellular networks were seen in the anterior ganglia of this specimen, but 
only a few in the posterior ganglia. X 1144. 
thetic trunks through two or three anterior white rami com- 
municantes and could pass downward in the trunks, giving off 
collaterals to several of the ganglia (Langley and Orbeli, ’11), or 
2) they might be the neuraxes or dendrites of commissural neurones 
with cell bodies located without the spinal cord. 
To exclude the possibility of cerebrospinal fibers reaching 
certain of the sympathetic ganglia a double operation was resorted 
to with the results reported below. 
