10 H. BERKELBACH VAN DER SPRENKEL 
easily distinguished from the sensory VII and nervus lateralis 
anterior by reason of the character of their fibers. The sequence 
of the different components of this root complex near its entrance 
is such that, in a transverse section, the motor V forms the mcst 
medial and ventral part of the complex. Next comes the sen- 
sory V, then the sensory VII, and most dorsal the nervus later- 
alis anterior. These roots shift gradually into the brain stem. 
The intramedullary course of the motor V is nearly horizontal 
and inclined in a caudal direction. Near the frontal pole of its 
nucleus it curves inward around the anterior secondary gusta- 
tory tract (fig. 6), and thus reaches its motor cells. The fibers— 
which are coarse and easily distinguished from the surrounding 
tracts—clearly form, two bundles, a frontal one arising from the 
frontal pole of the nucleus and a caudal one originating in its 
caudal part. I have not been able to decide whether these bun- 
dles remain thus separated during their further course or whether - 
they again mingle. The motor nucleus has an extremely ventral 
position (figs. 6 and 21), just mesially of the entrance of the motor 
VII root. The cells are large and have considerable dendritic 
outgrowths. Their position is entirely under and near the 
anterior secondary gustatory tract (fig. 6). In some places they 
form cellular columns which lie embedded among the associational 
systems. 
The motor V nucleus grows considerably smaller at about the 
middle of its length, thus exhibiting an isthmus which connects 
the two parts. A complete separation does not occur, though a 
division is thus clearly indicated. Such an arrangement of the 
motor V nucleus in two divisions has also been described in 
Tinea, which by the ventral position of the V nucleus (fig. 21) 
likewise exhibits a striking resemblance to Silurus. Both animals 
are characterized by a considerable development of the organs of 
taste. 
As was stated above, the sensory V8 has a position lateral to 
the corresponding motor root outside the medulla (fig. 5). Where 
the motor V curves around the anterior secondary gustatory 
8 The mesencephalic root of the V nerve could not be traced with certainty. 
