HuBER, Sympathetic Nervous System. 125 



ated. The degenerated fibers could be traced to the peri- 

 phery of the ciHary ganglion but not beyond the ganglion 

 cells, while the short ciliary nerves were never found degen- 

 erated. These experiments show very conclusively that these 

 medullated iibers ending in the ciliary ganglion, are the neur- 

 axes of neurons situated central to the point of section. The 

 non-degeneration of the short ciliary branches can only be ex- 

 plained by the fact that they are neu raxes of neurons, the 

 trophic center of which — the cell body and nucleus of the 

 neuron — are situated peripheral to the point of section, namely 

 in the ciliary ganglion. . Sections of the ganglion and nerves 

 of the normal side containing no degenerated fibers show 

 however that the fibers entering the ganglion from the third 

 nerve are very small medullated fibers, which can be likened to 

 the central fibers ending in other sympathetic ganglia. 



The fibers from the Vth nerve and those from the sympa- 

 thetic, forming the sensory and sympathetic roots do not end in 

 the ganglion, as may be seen from Langley and Anderson's (83) 

 and from Apolant's (86) work. 



Thus far my discussion has been largely concerned with 

 the larger ganglia of the chain and the cranial ganglia, into 

 which the small medullated nerves — central fibers — can be traced 

 without much difficulty. 



It must not however be supposed that such nerves end 

 only in these larger ganglia. All evidence we possess goes to 

 show that similar nerves end in all the sympathetic ganglia, 

 even the small terminal ganglia. Attention has already been 

 drawn to the fact that intra-capsular, peri-cellular baskets — the 

 endings of the small medullated, the central or pre-gangli- 

 onic fibers — are to be found in even the smallest peripheral 

 or terminal ganglia. This alone would justify the assump- 

 tion that pre-ganglionic fibers end in the peripheral ganglia. 

 It is however not difficult to show, (and especially in meth- 

 len-blue preparations) that small medullated fibers, larger than 

 the medullated fibers of the sympathetic system, enter the small 

 peripheral ganglia and terminate in end-baskets. We must as- 

 sume then that many of the nerve fibers of the white rami pass 



