Herrick, Zerminal Buds of Fishes. 125 
unspecialized visceral sensory fibers must be treated together 
within the brain, since they are so intimately intermingled that 
analysis has so far proven impossible. Accordingly, they are 
termed collectively the ‘‘communis system”’ of cranial nerve 
fibers. Peripherally this system is clearly divisible into two 
components, (1) the unspecialized visceral sensory which is 
doubtless much more ancient phylogenetically, and (2) the 
specialized, which for reasons to appear we may now term the 
gustatory component. This component may be divided topo- 
graphically into two divisions, one for taste buds within the 
mouth, and one for terminal buds in the outer skin. It is the 
latter division only, of course, with which we are here con- 
cerned. 
In fishes in which the terminal buds are not exceedingly 
numerous, as in the cod, their nerve fibers, though they enter 
the brain by the facial nerve, all turn back in the fasciculus 
communis to find their terminal nucleus in the vagal lobe, along 
with all other communis nerves of the body. But in the two 
groups of teleostean fishes in which these organs are most 
abundant (viz., the siluroids and cyprinoids) their nerve fibers 
find their terminal nucleus at the cephalic end of the fasc. com- 
munis in a special center, the lobus facialis. In both of these 
groups. all of the terminal buds are innervated from the genicu- 
late ganglion of the facialis root. Of this I am confident in the 
siluroids (HERRICK, ’o1). As to the cyprinoids, a rather 
cursory examination of sections of Carassius convinces me that 
the same is true, though my study of this type is not by any 
means exhaustive. 
The facial lobe of the siluroids was formerly known as the 
lobus trigemini, that of the cyprinoids as the lobus impar, in 
the former case the structure being paired, in the latter un- 
paired by the fusion in the dorso-median line of the lobes of 
the two sides. That the facial lobes in these two groups of 
fishes are really merely differentiated parts of the vagal lobes is 
manifest, not only from the nature of their peripheral connec- 
tions as indicated above, but still more evidently from their. in- 
ternal structure and secondary fiber connections. I can speak 
