262 Journal of Comparative Neurology. 



r. ophthalmicus profundus V in Amblystoma and the tadpole, 

 follows from the conclusions already discussed. The lateral and 

 mesial terminal branches of this nerve in Amblystoma innervate 

 an area which corresponds very closely to that innervated in 

 Rana by the medialis and lateralis narium of Gaupp ('97) and 

 the r. maxillaris. Within this area in Amblystoma the varia- 

 tion of the manner of branching is so great that it is impossi- 

 ble to establish any positive relation between the two forms in 

 this particular. It seems reasonable that the communicating 

 nerves of the general cutaneous component of the trigeminus 

 to the r. palatinus VII in Anura have their equivalents in the 

 two divisions of the ventral terminal branches of the r. oph- 

 thalmicus profundus in Amblystoma. The divergence of Am- 

 blystoma and Rana in this portion of the peripheral nervous 

 system is much greater, I believe, than has been generally 

 accredited ; and the application of anuran terminology to the 

 nerves of Urodela, as Boweks has applied it to Spelerpes, is 

 not only incorrect in this case but tends to obscure the actual 

 condition of relationship. 



J. — Minor Branches of the Trigeminus. 



The second and third branches of the r. ophthalmicus pro- 

 fundus in Amblystoma seem to be represented in Spelerpes by 

 a single nerve which is called " Va" by Bowers. The first 

 cutaneous branch of the r. mandibularis V of Amblystoma, 

 also, is represented in Spelerpes by a nerve which Bowers fig- 

 ures as "Vd." The latter nerve is probably represented in 

 Cryptobranchus by the nerve which McGregor describes as 

 penetrating the m. masseter and running "latero-caudad," 

 supplying the skin over the muscle mentioned and about the 

 angle of the jaw. McGregor says this nerve "may represent 

 Strong's ramus accessorius. " Such a relation, however, 

 seems wholly improbable, as shown by the above discussion of 

 the latter nerve. 



The anastomosis between the r. mandibularis V and the r. 

 alveolaris VII is not found by Bowers in Spelerpes. This 

 relation will be noticed particularly in connection with the r. 



