THE OLFACTORY NERVE OF LEPIDOSTEUS 129 



fairly compact central ganglion and one or more peripheral 

 ganglia persisting through all stages to the adult. 



Belogolowy ('12), who has followed the development of the 

 nervus terminalis in selachians, finds a ganglion to originate from 

 the olfactory placode (p. 8) in accord with my description above, 

 but would interpret this ganglion as representing in part a primi- 

 tive olfactory ganglion (p. 11). As far as the majority of the 

 embryological evidence goes, therefore, we may say that the 

 nervus terminalis is a component of the olfactor}^ (or primitive 

 olfactory) nerve, and this might very well be the case, although it 

 does not have the same central connection with the brain, as 

 Johnston ('13, p. 108) has very well said. As to the function of 

 the nervus terminalis, the present investigation has no new evi- 

 dence. As we ('10) previously stated for Amia, the circum- 

 stantial evidence leads one to ascribe a vasomotor function to it 

 in part. The disposition of the cells in Lepidosteus in more 

 compact central and diffuse peripheral ganglia allows of its falling 

 quite naturally into the morphological relations of the typical 

 autonomic system. It may be noted that Huber and Guild 

 ('13, p. 272) are somewhat inclined to assigning it to the sympa- 

 thetic system. 



I wish to acknowledge my indebtedness to the Ohio Academy 

 of Science for a grant from the AlacMillin Fund to defray the 

 expense of collecting material, and to my wife for helping with the 

 drawings. 



Little Rock, Arkanaaa, January 14, 1914 



THE JOURNAL OF COMPARATIVE NEUROLOGY, VOL. 24, NO. 2 



