NERVUS TERMINALIS: MAMMALS 43 



and anterior ethmoidal branches of the trigeminal nerve. As 

 already noted, there is a commingling of fibers of this nerve and 

 of the terminalis, and fibers of the trigeminus enter the bundles 

 which constitute the septal plexus of the terminalis. The intri- 

 cacy of this plexus made it impossible to follow any individual 

 fiber very far. Accordingly it was not possible to detennine 

 with certainty whether the free terminations of the septal mucosa 

 are from terminalis fibers or from the trigeminal nerve. 



Other fibers of larger size are also present in the mucosa. These 

 terminate on or near the septal glands, and show varicosities on 

 their finer twigs. They also enter into the plexus of the termi- 

 nalis to some extent. They are given off from the fifth nerve. 

 It is usually stated that the septal glands are innervated by the 

 trigeminus, and these fibers appear to be the ones by which 

 this is accomplished. 



2. The nervus terminalis of the beef 



The nervus tenninahs of the beef, as shown in figure 32, lies 

 median to the olfactory nerves, between the meninges and the 

 ventral brain surface. Running parallel with it are branches 

 of the anterior cerebral artery. In the specimen figured the 

 greater portion of the left nerve is a compact bundle, while the 

 right nerve is composed of several strands for the greater part of 

 its length. In the numerous brains examined there was con- 

 siderable variation in the relation of the different strands which 

 by their union form the main nerve bundle. This variation was 

 found not only when comparing one brain with another, but, as 

 just indicated, on comparing the two nerves of the same speci- 

 men. In some cases the strands were independent up to a 

 short distance from the forward margin of the hemispheres, in 

 other cases the bundle was formed much further caudad. 



For the greater part of its course the portion of the nerve pres- 

 ent in the specimens was covered by the meningeal membranes. 

 It emerges to the outer surface of the arachnoid coat at about 

 the point where the cerebral hemispheres begin to curve upward. 

 At this point the nerve was always compact in a single bundle. 



