454 F. W. CARPENTER 
are concentrically arranged about the capsules of the ganglion 
cells. From them a few centrally directed offshoots ending in 
knobsare traceable through the capsular walls to the surfaces of the 
underlying cell-bodies. That these extracapsular, nest-like struc- 
tures are not identical with the intracapsular end nets described 
above is clear from their position, and the inter-relations of their 
fibers, which do not appear to anastomose. They are doubtless 
bundles of preganglionic neurites on the way to their subcapsular 
terminations, the fibers given off to the cell-bodies being the ter- 
minal portions of such neurites. The latter, as has been said, 
end in knob-like swellings which are in contact with the ganglion 
cells. Such simple end organs may, in the human sphenopalatine 
ganglion, put the preganglionic neurites in communication with 
the postganglionic neurones. However, the foregoing results 
with methylene blue staining in the sheep raise the question if 
these knobs are not, in reality, the first varicosities of an intracap- 
sular end net, the remainder of which has not been differentiated 
by the Bielschowsky technique. 
SUMMARY 
The sphenopalatine, otic and submaxillary ganglia of the sheep 
contain multipolar cells with long, slender, frequently branched 
dendrites, which extend for considerable distances beyond the 
limits of the cell capsules. They resemble in these particulars the 
ordinary type of mammalian sympathetic cells. 
In the ciliary ganglion the only cells in which processes were 
clearly differentiated by methylene blue possessed each a single, 
heavy, branched dendrite. 
In all the cranial autonomic ganglia (ciliary, sphenopalatine, 
otic, submaxillary) the preganglionic neurites terminate on the 
cell-bodies of the postganglionic neurones in subcapsular, peri- 
cellular end nets of fine varicose fibrils. These endings are similar 
to those of preganglionic fibers in the vertebral and prevertebral 
ganglia of the sympathetic system. 
