GANGLIA OF RANA 477 
its cells ef origin lie in the posterior portion of branchial X. It 
is hardly probable that all of them lie in branchial X, so that 
until the embryological origin is worked out the problem of the 
exact morphology of branchial X! will have to be left as indicated 
above, i.e., that a part of the cells from which the second branchial 
nerve (6 of Strong’s plot) originates lie in the visceral ganglion and 
that the visceral ganglion represents the general visceral ganglion 
of such types as Menidia, Lepidosteus and Ameiurus, plus the 
representative, one or more, of branchial ganglia of these types. 
‘SUMMARY AND DISCUSSION 
1. The trigemino-facial complex of the frog in stages earlier 
than those studied by Strong corresponds in all essential details 
with his analysis except in the greater isolation of the profundus. 
This ganglion in earlier stages is much more isolated than in 
the earliest plot given in this paper, but even in that it stands out. 
rather distinctly, indicating its definite character which is lost 
by incorporation with the Gasserian. In other respects we con- 
firm Strong’s account of the V + VII ganglia in the frog. 
2. The glossopharyngo-vagal complex of ganglia, on the con- 
trary, if taken in the 8 mm. and 10 mm. stages, shows a much 
greater degree of simplicity and isolation of its various compo- 
nents than indicated by Strong and furthermore it is much more 
typical as compared with such types as Menidia, Ameiurus and 
Lepidosteus. 
3. The lateralis components in the glossopharyngo-vagal 
complex are represented by three more or less distinct ganglia. 
These are, (a) a lateralis [X situated on the root of the [X behind 
and at the level of the middle of the auditory vesicle dorso-ven- 
trally; (b) two lateralis X ganglia situated on the lateral surface 
of the cutaneous and general visceral ganglia respectively. Of 
these two ganglia, the dorsal lateralis is proximal and the ventral 
lateralis distal. The dorsal lateralis lies lateral to the jugular, or 
general cutaneous X, and gives rise to one nerve trunk which 
immediately after leaving the ganglion splits, giving rise to the 
most posterior ramus of Strong’s plot (dorsalis and medialis of 
