GANGLIA OF RANA 479 
the [IX + X complex. It is situated mesial to the dorso-lateral 
X and is consequently proximal of the visceral X. It is very 
large in the tadpole, much larger than in the Lepidosteus and 
Ameiurus at the same stage of growth and is much farther away 
from the medulla. In the 35 mm. tadpole, however, it migrates 
towards the medulla and les partly, though not wholly, in the 
jugular foramen. 
One nerve arises pure from this ganglion at this stage, the ramus 
auricularis; others arise also but run out in conjunction with 
other components and could not be accurately traced. So far as 
they could be followed their distribution agrees with Strong’s 
description. 
5. The visceral ganglia of the glossopharyngo-vagal complex 
lie in three well defined masses: (a) the glossopharyngeal; (b) 
the first or most anterior visceral, and (c) the second or most 
posterior visceral. 
(a) The glossopharyngeal ganglion occupies a position ventral 
to the auditory capsule with its long axis parallel to that of the 
body. The root is long and curves around behind the auditory 
capsule to enter the medulla along with those of X. It seems to 
be a pure visceral ganglion, although a cutaneous ramus, r. com- 
municans IX ad VII, passes out from the anterior end of the 
ganglion. Its fibers, however, can be traced past the ganglion 
and apparently enter jugular X, as described by Strong. It was 
not possible in the stages studied to recognize the special visceral 
or gustatory portion of this ganglion. ‘Two nerves, the ramus 
pharyngeus and the ramus laryngeus, arise from the anterior end 
of this ganglion. 
(b) The first visceral or branchial ganglion of X resembles 
in shape and position the glossopharyngeal. It lies under the 
posterior end of the auditory capsule with its long axis parallel 
to that of the body. Its posterior end is not, like that of IX, 
continued into a fibrous root but joins the cell mass of the second 
visceral X near its middle region. Gustatory cells could not be 
recognized in this ganglion in the stages studied. Two large 
nerves arise from this ganglion, the branchial 6 and 7 of Strong’s 
plot. 
