342 ELIZABETH CAROLINE CROSBY 
On the other hand, Elliot Smith (10) has considered the 
epistriatum to be of cortical origin. He believes that the effect- 
ing of olfacto-somatic correlations in the reptilian hemisphere 
and, particularly, the entrance of tactual fibers into the dorsal 
part of the hemisphere lead to the disturbance of the morpho- 
logical relations of the centers of the forebrain. He says ‘‘One 
curious manifestation of these disturbing influences is seen in 
the ingrowth (toward the lateral ventricle) of part of the over- 
growing pallium, forming a structure to which Edinger gave the 
name ‘epistriatum.’ The epistriatum is not a part of the striate 
body but is cortical in nature. Moreover it is not a morpholog- 
ical subdivision of the hemisphere which can be identified in 
other groups of vertebrates, as many anatomists believe. It 
is merely a peculiar adaptation of structure to meet the con- 
ditions favorable to the reptile;—namely the disturbing influ- 
ence of the recent admission of tactile impressions into the 
hemisphere.” 
In his 1915 paper, Johnston, following Edinger and Kappers 
(Kappers, 06, p. 9), suggests that the term ‘epistriatum’ be 
dropped, basing his suggestion on the facts that “the structure 
to which the term was first applied, does not appear as a special 
body or ridge in the turtle brain” and that “the author 
(Edinger) of the term uses it for at least three different bodies 
in the reptilian brain.”’ The conditions found in the forebrain of 
Alligator mississippiensis certainly support Johnston’s suggestion. 
In the anterior end of the hemisphere of the alligator several 
large cell masses are found in the basal portion of the lateral wall. 
1. There is a dorso-lateral area which includes a part or all 
of the ‘epistriatum’ as that term is used by some recent writers 
on the reptilian brain and is comparable with the dorsal ventric- 
ular ridge (Johnston, ’15) in turtles, though perhaps not exactly 
homologous with that area. 2. Below the dorso-lateral area, in 
a ventro-medial position, are two nuclei which belong to the cor- 
pus striatum of Johnston. The more dorsal large celled mass 
is the ventro-lateral, large celled nucleus of this description, 
comparable with Johnston’s nucleus lentiformis in the turtle. 
3. The more ventral small celled mass is the ventro-lateral, 
