CORONATE — PARAPHVLLINA. 549 



Genus PARAPHYLLINA Maas, 1903. 



Parajihyllina, ^L\As, 1903, Scyphomedusen der Siboga Expedition, Monog. 11, p. 6; 1907, Ergeb.und Fort sell ritte dcr Zool., Bd. 1, 

 p. 195. 



The type species is Parapliylltna intermedia Maas, 1903; I'lom the Malay Archipelago, 

 between 100 fathoms and tiie surtace. 



GENERIC CHARACTERS. 



Coronatae with 12 tentacles, 4 perradial marginal sense-organs, and 16 lappets. The 12 

 tentacles are interradial and adradial with reference to the stomach and lips. There are 

 4 pairs (8) of interradial gonads. The marginal sense-organs have a terminal lithocyst-sac 

 and a ventral bulb-like swelling, just beyond which is an eye with ectodermal lens and ecto- 

 dermal pigment. 



The short 4-sided throat-tube and flat disk-like bell resemble the condition noted in 

 Nausithoe, but in its 4 marginal sense-organs and 12 tentacles it recalls the condition seen in 

 Pcriphylla, with the important difference that in Periphylla the sense-organs are interradial, 

 whereas in Paraphyllina they are perradial in position. 



Maas places this genus in a new family which he calls the Paraphyllinidae. The characters 

 of this family are those of its only genus, Paruphyllirui. It is closely related to the fossil genus 

 Paraphyllites, Maas, 1 906 (Neuen Jahrbiich. fiir Mineralogie, Geol. und Paliiontol., Bd. 12, 

 p. 90, 4 fign.). This fossil medusa differs from recent Paraphyllina only in that its 8 gonads are 

 adradial and placed 45° apart, whereas in Paraphyllina they are grouped in pairs on both 

 sides of the 4 interradii. 



Paraphyllites distinctus is described by Maas from a well-preserved specimen. It is 



15 mm. wide and has a well-developed coronal furrow, 16 pedalia, 4 perradial marginal 



sense-organs, 12 tentacles, and 8 adradial gonads 45° apart. It is from the lithographic slate 



of Kelheim. The latest reference to this fossil is that of von Amnion, 1908, Geonostischen 



Jahrespeften, Jahrg. 19, p. 170. 



Paraphyllina intermedia Maas. 



Paraphyllina intermedia, Maas, 1903, Scyphomedusen der Sihoga Expedition, Monog. 1 1, p. 8, taf. 2, fign. 10-14; taf. 1 1, fig. 106, 

 Periphylla dodecahoUrycha, Lobianco, 1903, MItth. Zool. Sta. Neapel, Bd. 16, p. 219, taf. 7, fig. 3. See also Maas, 1904, Result. 

 Camp. Sci. Prince de Monaco, fasc. 28, p. 48, foot-note. 



Bell flatly rounded, 8 mm. high and 15 mm. wide, without pointed apex. The coronal 

 furrow around the exumbrella is deep and separates the smooth, central, flatly-rounded apex 

 of exumbrella from the marginal zone of lappets. Lappet-zone about as wide as radius of 

 central disk of exumbrella. The 16 pedalia of the marginal zone are rectangular with rounded 

 angles and are separated one from another by deep radiating furrows. These pedalia are 

 in the radii of the tentacles and sense-organs and alternate with the lappets. The 12 pedalia 

 of the tentacles are of uniform width, while the 4 pedalia of the marginal sense-organs are 

 only about half as wide as the former. The 16 marginal lappets are oval and bluntly pointed, 

 and the 8 lappets flanking the 4 sense-organs are somewhat narrower than the others. The 

 12 tapering, hollow tentacles are all of equal length and are not quite as long as the diameter 

 of the bell. 



The 4 marginal sense-organs are each covered by a hood-like fold ot the exumbrella. 

 Each sense-club contains a small terminal sac-shaped entodermal concretion. On the ventral 

 (subumbrella) side of the sense-club is a large ectodermal eye with a cup-like mass ot pig- 

 ment and a spherical lens. On the inner side of the eye and upon the ventral side of the 

 sense-club is a bulb-like swelling. Altogether the sense-clubs resemble those of Nausithoe, 

 but the eye is larger. 



The throat-tube is short and 4-sided and the mouth is a simple cruciform opening. There 

 are 4 interradial rows of gastric filaments. The coronal ring-canal gives rise to 16 peripheral 

 pouches in the radii of the sense-organs and tentacles, and these are put into communication 

 one with another by means of a marginal ring sinus, as in Nausithoe. 



The gonads resemble those of Palephyra and are intermediate in character between 

 those o{ Periphylla and those of Nausithoe. They consist of 4 pairs of bean-shaped or egg- 

 shaped sacs. These 8 sacs project from the subumbrella floor on both sides of the 4 inter- 

 radii near the sides of the throat-tube and centripetal to the ring-muscle. 



