ORGANS FOR SPERM-TRANSFER 287 
of the two is reversed, raises the question as to whether the 
males are adjusted, in habit, to these two kinds of females, so 
as to use the right set of sperm transfer organs for a left-handed 
female and vice versa, or not. 
The crucial experiments to determine whether males actually 
use the one stylet for right and the other for left-handed females 
have not yet been made. However, some facts and considerations 
make it improbable that a male is obliged to do so and indicate 
that a male may adjust his stylets so as to use either right or left 
on any form of female annulus, leaving the question still open as 
to what is the normal habit of the males with reference to the two 
forms of females. 
In the first place we found that though the two first stylets seem 
to be in the annulus they are never both firmlyinserted. One is 
fixed firmly by its tip while the other may be drawn away by a 
pair of forceps. Moreover the one that is inserted has its tip 
some # to 14 mm. in advance of the other and its base is locked 
against the base of the other, diagonally, the abdomen being ad- 
vanced more on one side than the other. 
Observations showed that not only were there cases of the right 
stylet in the left annulus but of right stylet in the right annulus 
and of left stylet in left annulus and of left styletin right annulus. 
Whether in these cases the sperm was actually transferred was, 
unfortunately, not made out. It is possible that a male may in- 
sert one stylet and afterwards the other till finally the actual 
sperm transfer takes place with some more definite reference to 
the symmetry of the annulus than the above observations would 
indicate. 
That there is any alteration in the advance of the stylets was 
not made out, but there is often an alteration in the use of the fifth 
leg, right and left. At any one time many males will be found 
with the left and others with the right leg crossed, but continuous 
observations show that the male will change from right to left 
in difficult cases especially, till a better adjustment is obtained. 
It was at first thought that there was a relation between the 
fifth leg and the advance and use of the first stylet so that these 
were on the same side, that is, the stylet being advanced by the 
JOURNAL OF MORPHOLOGY, VOL. 22, No. 2 
