334 GILMAN A. DREW 
expulsion of a single jet of water. The male seemed to reach the 
female before she had time to move much, although she has given 
me the impression of attempting to dodge as if frightened. The 
two animals become attached head to head with their arms inter- 
mingled, each grasping the other (fig. 2). Then as in the other 
method, the male sweeps his left ventral arm past the end of the 
funnel and grasps the bundle of spermatophores. These are im- 
mediately thrust between the ventral armsof the female and held 
there for a few seconds. The animals then separate and exami- 
nation has shown fresh sperm reservoirs attached to the receiv- 
ing depression on the buccal membrane of the female. The empty 
cases of the ejaculated spermatophores may be held between the 
arms several minutes but they are finally dropped. Here, as in 
the other method of copulation, only the sperm reservoirs are 
retained for any length of time. 
The spermatophores begin to ejaculate immediately after leav- 
ing the penis and the whole process is completed in a very few 
seconds. Pulling the filament attached to the ejaculatory end of 
a spermatophore is all that is needed to start its ejaculation. As 
the ejaculatory end of the spermatophore leaves the penis last and, 
as the spermatophores in the penis and the spermatophoric sac 
are imbedded in a viscid secretion, there is every reason to believe 
that the pull given the spermatophores by the arm with which they 
are grasped, when this arm starts to transfer them from the penis 
to the mantle chamber or to the buccal membrane, is sufficient to 
start ejaculation. The arm carries the spermatophores into the 
position necessary for the attachment of the sperm reservoirs 
while they are ejaculating and holds them there until the ejacu- 
lation is complete and the reservoirs are attached. 
The structure of the spermatophores and the mechanics of 
ejaculation which lead to the attachment of the reservoirs will 
be treated in another paper. It should, however, beunderstood 
that the spermatophores are never attached as such, but they 
ejaculate and the sperm reservoirs are attached. As the reser- 
voirs are attached by cement carried inside the spermatophores 
and liberated by the ejaculation, they may be stuck anywhere. 
The sperm slowly escape from these reservoirs and may then 
