SOME PROBLEMS OF COELENTERATE ONTOGENY 501 
(09, p. 165), whose detailed experiments on differential staining 
have surpassed my own, was perplexed as to this condition. 
“At the end of the growth period, the nuclear reticulum shows so 
little affinity for basic stains that there appears to be, so far as 
this test shows, no chromatin present in the entire nucleus. I can 
suggest no explanation for this peculiar condition of the chromatin 
at this period, but it is normal and characteristic of this stage.”’ 
I am now convinced that there is here a chief ground for my fail- 
ure to distinguish certain phases of maturation, and my subse- 
quent error in the assumption of their possible suppression or 
modification in certain cases. 
3. Amuatosis. Concerning a further problem, that of amitosis, 
I amin doubt so far as Pennaria is concerned, even as at the time 
of my previous work. My chief grounds for this view are the facts 
first cited, and those of the multivesiculate aspects of the nucleus 
during cell proliferation. And here again Smallwood and G. T. 
Hargitt (09), and iater Beckwith (09), all confirm my basis of 
facts. They find in these vesiculate nuclear conditions essentially 
the same results which are described in my accounts. Without 
exception their interpretations differ from mine. To them these 
facts are believed to be obscure phases, chiefly telophases, of 
mitosis. While I freely admit the force of their contentions, 
there are still good reasons for maintaining the plausibility of 
my own views and interpretations. This is especially the case 
concerning Hudendrium. Here there seemed to be clear and posi- 
tive examples of amitosis, as shown in fig. 238, a and b, plate 
15 (04a). It may not be amiss to state here that all these exam- 
ples of amitosis occurred in association with those ‘nuclear nests’ 
so intimately involved in the syncytial phase of development con- 
cerned in entoderm formation. The conditions are somewhat 
different in Pennaria, yet sufficiently similar to lead one to anti- 
cipate similar processes, and these appeared probable in the vesi- 
culate ‘nuclear nests’ mentioned above. But in no case were 
there found the specific and positive examples figured in the case 
of Eudendrium. The same must also be said of Clava. But fur- 
ther discussion of this will be reserved for a later section. 
