588 VICTOR E. SHELFORD 
2. Relation of the behavior to habitat and associated forms 
There is the greatest difference in the behavior of the different 
species. I have as yet been unable to study this critically, but 
it is at least a very promising field. 
a. Behavior and habitat. As we have noted, C. sexguttata has 
for example, various peculiarities of behavior which are related 
to the forest conditions in which it lives, which are not possessed 
by other forms. As we noted, when it is frightened from a rock 
or bare place, it frequently alights on the leaves of a low tree or 
bush and crawls under the bark of trees for the night, or even 
to hide when pursued. None of the other species which I have 
studied behave in such a manner. I have never seen C. tranque- 
barica crawl under objects when pursued. It does not alight on 
the green leaves of trees or shrubs when they are present. It 
excavates burrows instead of crawling under objects. The be- 
havior of these two species is correlated with the general environ- 
mental conditions. 
b. Inter-physiology, and inter-psychology. Tarde (’03) has 
recently written an article on inter-psychology—the psychology 
of the relations of individuals of the same species (man). To 
this should be added the behavior between different species, while 
acting or living together as one. He suggests that the social 
psychology of man may be traced to the inter-psychology and 
physiology of the lower animals. If this is true, then we can 
be more certain that the inter-psychology of the higher forms has 
developed from the inter-physiology of the lower forms (Craig, 
08 [2]). 
I have looked for the inter-physiological manifestations in 
these beetles, but have found none striking except the mating 
instincts. There is little or no social life. I have found animals 
belonging to totally unrelated species attempting to copulate 
in some cases where the two are dissimilar. 
It seems quite evident from my observations that the more 
marked phases of the behavior of the tiger beetles arise not from 
inter-physiology, but from relations to the species which are quite 
