744 THOS. H. MONTGOMERY, JR. 
But this is unusual, for in the series of later prophases of figs. 
67 to 78 no trace of it wasnoticed. It is when the autosomes have 
shortened and their surfaces become nearly smooth that this 
split is prominent for the first time. The gemini of figs. 79, 80 
show it prominently but in each case upon only one univalent. 
It is fairly conspicuous along the larger geminus of fig. 82; in 
fig. 84 it is seen along two of the gemini; the reason why it is 
not seen upon all at this stage is probably because it is visible 
only when the flattened surface of a univalent is turned towards 
the observer. During the first maturation metaphase this split 
can always be seen when the gemini lie in the proper position and 
when they are sufficiently destained (figs. 86-88, 93-94). 
During the strepsinema stage and the consequent prophases ~ 
the nuclear linin threads increase in number, as shown in figs. 
42 to 93, and there are good indications that they are outgrowths 
of the autosomes. In the later prophases these become replaced 
by chains of fine globules (figs. 78-85). The latter appear to 
emanate from the autosomes, as though they were droplets of 
fluid pressed out by these during their process of condensation; 
and it may be that such droplets pass out along the linin threads, 
for this would explain their catenulated arrangement. This 
appears to be the only act of intranuclear substance emission by 
the autosomes, for in earlier periods the karyolymph appears 
quite clear. 
The history of the autosomes during the maturation mitoses 
has been so fully described by me before (’01, ’06, 710) that it 
needs but brief mention here. The six bivalent autosomes become 
arranged in the equatorial plane of the first maturation spindle 
with their long axes parallel to the latter (figs. 87-89; 92-94). 
All the chromosomes are drawn in fig. 89. Their longitudinal 
splits are also in the line of the spindle, and on polar views of the 
equatorial plate (figs. 90, 91) this split is often evident as an inden- 
tation upon the autosome. The transverse constriction of each 
geminus, which is the point of persisting conjugation of two uni- 
valents, lies in the equator and the autosomes divide along it 
(figs. 94, 95). This first division is then reductional, and gives: 
to each second spermatocyte six univalent autosomes. As the 
