SPERMATOGENESIS OF EUSCHISTUS 781 
mitochondria, consequently there is a variable and irregular 
portioning of them to the daughter cells, and it is a matter of 
chance how much mitochondrial substance goes into each of the 
latter. 
Fig. 101 shows them on a polar view of the equatorial plane 
of a second spermatocyte. Fig. 102 shows all the threads (12) 
of somewhat more than one hemisphere, and fig. 103 all (10) 
of one hemisphere of the same mitosis; in these metaphases most 
of the mitochondria are shorter than in the preceding division 
because most of them. had been divided then, but they lie quite 
as irregularly in the cell. Succeeding anaphases are exhibited 
in figs. 104, 107-109, illustrating that here the phenomena are 
essentially of the same kind as in the preceding mitosis, namely, 
those mitochondria that happen to lie across the eyuator become 
broken there by the cellular constriction while the others are 
not divided. In figs. 104 and 109 the mitochondria of only one 
hemisphere are drawn, but in figs. 107 and 108 all the mitochon- 
dria of the cell. ‘Though they are irregularly divided by the matu- 
ration divisions their relative amounts in the spermatids do not 
seem to be greatly different; the evidence is that each long mito- 
chondrial thread becomes divided in the second maturation mito- 
sis if not in the first. 
Curious protuberances of the spermatocytes are produced 
that would seem to indicate pseudopodial movements of the cell 
body during the maturation divisions. These are shown in 
figs. 88, 89, 92-95, and since all of these were drawn from cells 
floating free in the testicular cavity (except fig. 85) these cell 
processes are not due to the pressure of other cells. Similar 
protuberances are found upon second spermatocytes (figs. 102, 
104, 107); and the left hand one of fig. 104 looks as though it 
might have persisted since the stage of fig. 94. Frequently mito- 
chondria extend into such protuberances, but not always; there- 
fore they cannot be considered the producers of them. 
Towards the close of the second maturation mitosis the 
mitochondria of each spermatid commence to fuse together (figs. 
110, 111), forming an irregular mantle around the connective 
fibers of the spindle. Then those of each spermatid give rise 
JOURNAL OF MORPHOLOGY, VOL. 22, No. 3 
