504 A. H. STURTEVANT 
ing these two factors have these regions of a red or reddish color, 
so that the Brown Leghorn, and probably also the Columbian 
Wyandotte, must carry a factor for red, R. The factors J and 
N do not completely inhibit R, since most of my [-bearing birds 
show traces of brown here and there, and all the white necked 
males are very ‘brassy’ (yellowish). Both these characters some- 
times appear in pure Columbian Wyandottes. Apparently there 
is also a Leghorn pattern factor, L,2 causing black breast and 
black on the wing coverts in the male, and black stippling and 
salmon breast in the female, the latter effect appearing even 
in the presence of J. The factor LZ is hypostatic to another pat- 
tern factor, P, which is carried by the Columbian Wyandotte, 
and which inhibits all the colors just mentioned, as caused by 
L, leaving the color of the part dependent upon F and its inhib- 
itors. But one dose does not completely inhibit the stippling of 
the female. 
An alternative view, equally as satisfactory, I think, is that 
there is no inhibitor P, but that the Wyandotte has no L, and 
that the absence of this factor is dominant to its presence, 
heterozygous females being distinguishable by the stippling. On 
the first view L is probably present in all my birds. The con- 
stitution of the various types would then be as follows: 
Columbian Wyandotte INRPL 
Brown Leghorn inRpL 
Grayene INRPL or nRPL 
Red inRPL 
Silver gray INRpL, or InRpL 
Brown duckwing iINRpL 
One other combination is possible—iNRPL. This should 
give a bird with Columbian pattern, white or straw neck, and red 
body. It is possible that such would have appeared had more 
birds been raised, but I know of no variety having any similar 
color combination, and_have never observed it in a cross-bred 
2This is probably one of the components of the J (Jungle pattern factor) of 
Davenport (09). 
