506 A.“H. STURTEVANT 
IV. Gray & (gametes) INPM INpM InPM InpMiNPMiNpM inPM inpM 
Brown Leghorn @ inpM inpm 
INPM inpM —gray 
INpM inpM —duckwing 7 
InPM inpM —grayo 
InpM inpM —duckwing @ (silver gray) 
iNPM inpM —red & (white-necked?) 
iNpM inpM ~—duckwing @ 
inPM inpM —red& 
inpM inpM —brown oJ (not seen) 
INPM inpm —gray 9 
INpM inpm —silver gray 9 
InPM inpm —gray 9? 
InpM inpm —silver gray ? 
iNPM inpm —red @ (white-necked?) 
iNpM inpm —brown duckwing 9? 
inPM inpm —red 9° 
inpM inpm —brown 2 
OTHER EXPERIMENTS DEALING WITH THE SAME COLORS 
Bateson (’02, ’09) and Punnett (’05) have given some facts 
regarding the duckwing color. When the Brown Leghorn was 
crossed with the White Dorking or White Leghorn, they obtained 
in F, some silver gray females. Two of these mated to a pure 
Silver Gray Dorking male gave only silver grays, and of these, 
four females to a male gave only silver grays. From these facts 
Bateson (’09) infers that the replaced red and yellow of the 
Brown Leghorn probably depends upon a separate factor, which 
his white breeds lacked. It seems to me more probable that 
this factor, which I have called R, was present in all three 
breeds, and that the two white breeds carried also the factor 
I. Any F, female showing the silver gray color would then be 
as pure for J as a pure Dorking, the factor being sex-linked, 
which explains why they had no trouble in getting a dominant 
F, to breed true. 
Mr. T. Reid Parrish, a Columbian Wyandotte breeder, has 
published in advertising circulars and in poultry journals (e.g., 
Parrish, ’11) detailed accounts of how he originated a strain of 
Columbian Wyandottes (probably not the one used in my ex- 
periments). According to this account he used Light Brahma 
