SEX-LINKAGE IN FOWLS 507 
females with a White Wyandotte male. The Light Brahma has 
exactly the color of the Columbian Wyandotte. The breed 
seems to have been brought from the Orient in something like 
its present form, so that its history as to the origin of its color 
- must probably remain a matter of conjecture. The White Wyan- 
dotte was derived directly from the Silver Laced Wyandotte, 
and is still, or was comparatively recently, a not uncommon 
sport from that variety (see McGrew, ’01, and poultry literature 
generally). This would seem to indicate that it is a recessive 
white, probably due to the dropping out of a color producer. 
Mr. Parrish’s statements support this view, as he says he obtained, 
in the F; generation from his cross, silver laced, barred, and 
Columbian birds—apparently no whites. These F, silvers he says 
were not typical, some of them having nearly white breasts, “yet 
showing a trace of lacing throughout the plumage.” This sounds 
as though they were much like the birds I obtained in my cross: 
between silvers and Columbians (Sturtevant, 711). From the 
result of that cross it would appear that silver is incompletely 
hypostatic to Columbian. Mr. Parrish says of his F, barred 
birds mentioned above that they ‘‘showed stronger Brahma 
markings than the silvers, but there was unmistakable barring 
throughout the plumage, being especially noticeable in tail and 
wing, some specimens showing barring in every section.” This 
is a most interesting statement, in view of the work of Spillman 
(09), Goodale (’09, 710), Pearl and Surface (10), and Daven- 
port (706, ’09) on barring. In this connection it is worth noting 
that occasionally a few barred feathers occur in pure Columbian 
Wyandottes, especially in the tail coverts of young males, and that 
one of my F, males (a duckwing) in the experiment described 
above has some barring in his hackle. 
Mr. Parrish states that he mated his F,; Columbians with 
White Wyandottes, reciprocally. From Columbian female he 
obtained the same threeclasses as in F,, but we are not told 
whether or not whites appeared. The mating with Coltimbian 
male gave whites and Columbians, but he doesn’t say what else, 
if anything. 
